As another comment mentioned, papers get revised during review, usually in response to reviewer comments. Also, some journals (not sure about Nature) do not allow authors to "backport" revisions made in response to reviewer comments to preprints; I guess they view the review process as part of their "value add".
Its quite common to revise papers. For example, they might have uploaded to arxiv in order to submit to a conference. Later, they revised and submitted to Nature.